About the Authors

Xin Yin

Contributed equally to this work with: Xin Yin, Guohua Deng, Xianying Zeng

Roles Investigation, Methodology

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Guohua Deng

Contributed equally to this work with: Xin Yin, Guohua Deng, Xianying Zeng

Roles Funding acquisition, Investigation, Methodology, Validation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Xianying Zeng

Contributed equally to this work with: Xin Yin, Guohua Deng, Xianying Zeng

Roles Formal analysis, Investigation, Methodology, Validation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Pengfei Cui

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Yujie Hou

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Yanjing Liu

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Jingzhen Fang

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Shuxin Pan

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Dongxue Wang

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Xiaohan Chen

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Yaping Zhang

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Xiurong Wang

Roles Resources

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Guobin Tian

Roles Funding acquisition, Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Yanbing Li

Roles Funding acquisition, Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Yan Chen

Roles Funding acquisition, Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Liling Liu

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Yasuo Suzuki

Roles Resources

Affiliation College of Life and Health Sciences, Chubu University, Aichi, Japan

Yuntao Guan

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Chengjun Li

Roles Investigation

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Jianzhong Shi

Roles Conceptualization, Formal analysis, Investigation, Methodology, Validation, Writing – original draft

shijianzhong@caas.cn (JS); chenhualan@caas.cn (HC)

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Hualan Chen

Roles Conceptualization, Formal analysis, Funding acquisition, Methodology, Project administration, Supervision, Validation, Writing – original draft, Writing – review & editing

shijianzhong@caas.cn (JS); chenhualan@caas.cn (HC)

Affiliation State Key Laboratory of Veterinary Biotechnology, Harbin Veterinary Research Institute, CAAS, Harbin, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.