About the Authors

Elsa Ghirardini

Contributed equally to this work with: Elsa Ghirardini, Elena Restelli

Roles Investigation, Writing – original draft

Current address: Consiglio Nazionale delle Ricerche Institute of Neuroscience, Pisa, Italy

Affiliation Laboratory of Pharmacology and Brain Pathology, Humanitas Clinical and Research Center, Rozzano—Milan, Italy

Elena Restelli

Contributed equally to this work with: Elsa Ghirardini, Elena Restelli

Roles Investigation, Writing – review & editing

Affiliation Department of Neuroscience, Istituto di Ricerche Farmacologiche Mario Negri IRCCS, Milan, Italy

Raffaella Morini

Roles Investigation

Affiliation Laboratory of Pharmacology and Brain Pathology, Humanitas Clinical and Research Center, Rozzano—Milan, Italy

Ilaria Bertani

Roles Investigation

Affiliation Department of Neuroscience, Istituto di Ricerche Farmacologiche Mario Negri IRCCS, Milan, Italy

Davide Ortolan

Roles Investigation

Current address: Unit on Ocular and Stem Cell Translational Research, National Eye Institute, NIH, Bethesda, Maryland, United States of America

Affiliation Department of Neuroscience, Istituto di Ricerche Farmacologiche Mario Negri IRCCS, Milan, Italy

Fabio Perrucci

Roles Investigation

Affiliation Laboratory of Pharmacology and Brain Pathology, Humanitas Clinical and Research Center, Rozzano—Milan, Italy

Davide Pozzi

Roles Investigation

Affiliation Laboratory of Pharmacology and Brain Pathology, Humanitas Clinical and Research Center, Rozzano—Milan, Italy

Michela Matteoli

Roles Conceptualization, Funding acquisition, Supervision, Writing – review & editing

michela.matteoli@hunimed.eu (MM); roberto.chiesa@marionegri.it (RC)

Affiliations Laboratory of Pharmacology and Brain Pathology, Humanitas Clinical and Research Center, Rozzano—Milan, Italy, Consiglio Nazionale delle Ricerche Institute of Neuroscience, Milan, Italy

Roberto Chiesa

Roles Conceptualization, Funding acquisition, Supervision, Writing – original draft, Writing – review & editing

michela.matteoli@hunimed.eu (MM); roberto.chiesa@marionegri.it (RC)

Affiliation Department of Neuroscience, Istituto di Ricerche Farmacologiche Mario Negri IRCCS, Milan, Italy

Competing Interests

The authors have declared that no competing interests exist.