About the Authors

Xiaojian Zhang

Contributed equally to this work with: Xiaojian Zhang, Hailiang Sun, Fred L. Cunningham, Lei Li

Roles Writing – original draft, Writing – review & editing

Affiliation Department of Basic Sciences, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America

Hailiang Sun

Contributed equally to this work with: Xiaojian Zhang, Hailiang Sun, Fred L. Cunningham, Lei Li

Roles Writing – original draft, Writing – review & editing

Affiliations Department of Basic Sciences, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America, College of Veterinary Medicine, South China Agricultural University, Guangzhou, Guangdong, China

Fred L. Cunningham

Contributed equally to this work with: Xiaojian Zhang, Hailiang Sun, Fred L. Cunningham, Lei Li

Roles Writing – review & editing

Affiliation Mississippi Field Station, National Wildlife Research Center, Wildlife Services, Animal and Plant Health Inspection Service, United States Department of Agriculture, Starkville, Mississippi State, Mississippi, United States of America

Lei Li

Contributed equally to this work with: Xiaojian Zhang, Hailiang Sun, Fred L. Cunningham, Lei Li

Roles Writing – review & editing

Affiliation Department of Basic Sciences, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America

Katie Hanson-Dorr

Roles Writing – review & editing

Affiliation Mississippi Field Station, National Wildlife Research Center, Wildlife Services, Animal and Plant Health Inspection Service, United States Department of Agriculture, Starkville, Mississippi State, Mississippi, United States of America

Matthew W. Hopken

Roles Writing – review & editing

Affiliations National Wildlife Research Center, Wildlife Services, Animal and Plant Health Inspection Service, United States Department of Agriculture, Fort Collins, Colorado, United States of America, Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado State, Colorado, United States of America

Jim Cooley

Roles Writing – review & editing

Affiliation Department of Pathobiology and Population Medicine, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America

Li-Ping Long

Roles Writing – review & editing

Affiliation Department of Basic Sciences, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America

John A. Baroch

Roles Writing – review & editing

Affiliation National Wildlife Research Center, Wildlife Services, Animal and Plant Health Inspection Service, United States Department of Agriculture, Fort Collins, Colorado, United States of America

Tao Li

Roles Writing – review & editing

Affiliation Viral Diseases Branch, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Brandon S. Schmit

Roles Writing – review & editing

Affiliation National Wildlife Research Center, Wildlife Services, Animal and Plant Health Inspection Service, United States Department of Agriculture, Fort Collins, Colorado, United States of America

Xiaoxu Lin

Roles Writing – review & editing

Affiliation Viral Diseases Branch, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Alicia K. Olivier

Roles Writing – review & editing

Affiliation Department of Pathobiology and Population Medicine, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America

Richard G. Jarman

Roles Writing – review & editing

Affiliation Viral Diseases Branch, Walter Reed Army Institute of Research, Silver Spring, Maryland, United States of America

Thomas J. DeLiberto

Roles Writing – review & editing

Thomas.J.DeLiberto@aphis.usda.gov (TD); wan@cvm.msstate.edu (XFW)

Affiliation National Wildlife Research Center, Wildlife Services, Animal and Plant Health Inspection Service, United States Department of Agriculture, Fort Collins, Colorado, United States of America

Xiu-Feng Wan

Roles Writing – review & editing

Thomas.J.DeLiberto@aphis.usda.gov (TD); wan@cvm.msstate.edu (XFW)

Affiliation Department of Basic Sciences, College of Veterinary Medicine, Mississippi State University, Starkville, Mississippi State, Mississippi, United States of America

Competing Interests

The authors have declared that no competing interests exist.