About the Authors

William H. Palmer

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Project administration, Visualization, Writing – original draft, Writing – review & editing

W.H.Palmer@sms.ed.ac.uk

Affiliation Institute of Evolutionary Biology University of Edinburgh, Charlotte Auerbach Road, Edinburgh, United Kingdom

Nathan C. Medd

Roles Investigation

Affiliation Institute of Evolutionary Biology University of Edinburgh, Charlotte Auerbach Road, Edinburgh, United Kingdom

Philippa M. Beard

Roles Methodology, Resources

Current address: The Pirbright Institute, Pirbright, Surrey, United Kingdom

Affiliation The Roslin Institute and The Royal (Dick) School of Veterinary Studies, University of Edinburgh, Midlothian, United Kingdom

Darren J. Obbard

Roles Conceptualization, Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

Affiliations Institute of Evolutionary Biology University of Edinburgh, Charlotte Auerbach Road, Edinburgh, United Kingdom, Centre for Infection, Evolution and Immunity, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.