About the Authors

Chikako Ono

Contributed equally to this work with: Chikako Ono, Takasuke Fukuhara

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Takasuke Fukuhara

Contributed equally to this work with: Chikako Ono, Takasuke Fukuhara

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Daisuke Motooka

Affiliation Department of Infection Metagenomics, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Shota Nakamura

Affiliation Department of Infection Metagenomics, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Daisuke Okuzaki

Affiliation DNA-Chip Developmental Center for Infectious Diseases, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Satomi Yamamoto

Current address: Laboratory of Veterinary Microbiology, School of Veterinary Medicine, Kitasato University, Aomori, Japan

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Tomokazu Tamura

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Hiroyuki Mori

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Asuka Sato

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Kentaro Uemura

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Yuzy Fauzyah

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Takeshi Kurihara

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Takahiro Suda

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Osaka University, Osaka, Japan

Akira Nishio

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Osaka University, Osaka, Japan

Su Su Hmwe

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Toru Okamoto

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Tomohide Tatsumi

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Osaka University, Osaka, Japan

Tetsuo Takehara

Affiliation Department of Gastroenterology and Hepatology, Graduate School of Medicine, Osaka University, Osaka, Japan

Kazuaki Chayama

Affiliation Department of Medicine and Molecular Science, Hiroshima University School of Medicine, Hiroshima, Japan

Takaji Wakita

Affiliation Department of Virology II, National Institute of Infectious Diseases, Tokyo, Japan

Kazuhiko Koike

Affiliation Department of Gastroenterology, Graduate School of Medicine, The University of Tokyo, Tokyo, Japan

Yoshiharu Matsuura

matsuura@biken.osaka-u.ac.jp

Affiliation Department of Molecular Virology, Research Institute for Microbial Diseases, Osaka University, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: CO TF KK YM. Formal analysis: CO DM SN DO. Investigation: CO TF SY TTam HM AS KU YF TK TS AN TO. Project administration: CO TF. Resources: TS AN SSH TTat TTak KC TW. Writing – original draft: CO TF DM SN DO YM.