About the Authors

Nelson D. Glennie

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Susan W. Volk

Affiliation Department of Clinical Studies-Philadelphia, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Phillip Scott

pscott@vet.upenn.edu

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: NDG PS. Formal analysis: NDG. Funding acquisition: PS. Investigation: NDG SWV. Methodology: NDG SWV. Project administration: NDG. Resources: PS. Supervision: PS. Validation: NDG. Visualization: NDG. Writing – original draft: NDG. Writing – review & editing: PS SWV.