About the Authors

Cierra N. Casson

Current address: Immunology Program, Benaroya Research Institute at Virginia Mason, Seattle, Washington, United States of America

‡These authors contributed equally to this work and are listed in alphabetical order.

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jessica L. Doerner

‡These authors contributed equally to this work and are listed in alphabetical order.

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alan M. Copenhaver

Current address: Department of Respiratory, Inflammation and Autoimmunity, MedImmune LLC, Gaithersburg, Maryland, United States of America

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jasmine Ramirez

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Alicia M. Holmgren

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Mark A. Boyer

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Ingharan J. Siddarthan

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sara H. Rouhanifard

Affiliation Department of Bioengineering, School of Engineering and Applied Science, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Arjun Raj

Affiliation Department of Bioengineering, School of Engineering and Applied Science, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Sunny Shin

sunshin@mail.med.upenn.edu

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: one of the authors, Alan M. Copenhaver, is employed by a commercial company, MedImmune LLC. This does not alter our adherence to all PLOS policies on sharing data and materials.

Author Contributions

Conceptualization: CNC JLD AMC SS. Formal analysis: CNC JLD AMC JR AMH MAB IJS SHR AR SS. Funding acquisition: CNC JLD AMC AR SS. Investigation: CNC JLD AMC JR AMH MAB IJS SHR. Methodology: CNC JLD AMC AMH SHR AR SS. Project administration: CNC JLD AMC SS. Resources: AR SS. Software: SHR AR. Supervision: AR SS. Validation: CNC JLD AMC JR AMH MAB IJS SHR AR SS. Visualization: CNC JLD AMC JR AMH MAB SHR SS. Writing – original draft: CNC JLD AMC SS. Writing – review & editing: CNC JLD AMC AMH SHR AR SS.