About the Authors

Alec J. Hirsch

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Jessica L. Smith

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Nicole N. Haese

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Rebecca M. Broeckel

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Christopher J. Parkins

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Craig Kreklywich

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Victor R. DeFilippis

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Michael Denton

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

Patricia P. Smith

Affiliation Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America

William B. Messer

Affiliation Department of Molecular Microbiology and Immunology, Oregon Health and Science University, Portland, Oregon, United States of America

Lois M. A. Colgin

Affiliation Pathology Services Unit, Division of Comparative Medicine, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Rebecca M. Ducore

Affiliation Pathology Services Unit, Division of Comparative Medicine, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Peta L. Grigsby

Affiliation Division of Reproductive & Developmental Sciences, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Jon D. Hennebold

Affiliations Division of Reproductive & Developmental Sciences, Oregon National Primate Research Center, Beaverton, Oregon, United States of America, Department of Obstetrics & Gynecology, Oregon Health and Science University, Portland, Oregon, United States of America

Tonya Swanson

Affiliation Division of Pathobiology and Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Alfred W. Legasse

Affiliation Division of Pathobiology and Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Michael K. Axthelm

Affiliation Division of Pathobiology and Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Rhonda MacAllister

Affiliation Clinical Medicine Unit, Division of Comparative Medicine, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Clayton A. Wiley

Affiliation UPMC Presbyterian Hospital, Division of Neuropathology, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Jay A. Nelson

Affiliations Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America, Division of Pathobiology and Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Daniel N. Streblow

streblow@ohsu.edu

Affiliations Vaccine and Gene Therapy Institute, Oregon Health and Science University, Beaverton, Oregon, United States of America, Division of Pathobiology and Immunology, Oregon National Primate Research Center, Beaverton, Oregon, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: AJH MKA PLG JDH JAN DNS. Data curation: AJH DNS. Funding acquisition: AJH DNS. Investigation: AJH JLS NNH RMB CJP CK VRD MD PPS WBM LMAC RMD PLG JDH TS AWL RM CAW DNS. Methodology: AJH JLS CJP CK AWL DNS. Project administration: AJH DNS. Resources: AJH JLS CAW JDH JAN DNS. Supervision: AJH JDH MA RM CAW JAN DNS. Validation: AJH DNS. Visualization: AJH JLS VRD CAW DNS. Writing – original draft: AJH VRD CAW DNS. Writing – review & editing: AJH JLS NNH RMB CJP VRD PPS WBM LMAC RMD PLG JDH RM CAW JAN DNS.