About the Authors

Sama Adnan

Contributed equally to this work with: Sama Adnan, R. Keith Reeves

Affiliations Yerkes National Primate Research Center, Emory University, Atlanta GA, United States of America, New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America

R. Keith Reeves

Contributed equally to this work with: Sama Adnan, R. Keith Reeves

Affiliations New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America, Center for Virology and Vaccine Research, Beth Israel Deaconess Medical Center, Boston, MA, United States of America

Jacqueline Gillis

Affiliation New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America

Fay E. Wong

Affiliation New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America

Yi Yu

Affiliation New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America

Jeremy V. Camp

Affiliation Department of Microbiology, Immunology and Parasitology, Louisiana State University Health Sciences Center, New Orleans, Louisiana, United States of America

Qingsheng Li

Affiliation Nebraska Center for Virology and School of Biological Sciences, University of Nebraska-Lincoln, Lincoln, Nebraska, United States of America

Michelle Connole

Affiliation New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America

Yuan Li

Affiliation AIDS and Cancer Virus Program, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, Maryland, United States of America

Michael Piatak Jr

† Deceased.

Affiliation AIDS and Cancer Virus Program, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, Maryland, United States of America

Jeffrey D. Lifson

Affiliation AIDS and Cancer Virus Program, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, Maryland, United States of America

Wenjun Li

Affiliation Department of Medicine, University of Massachusetts Medical School, Worcester, MA, United States of America

Brandon F. Keele

Affiliation AIDS and Cancer Virus Program, Leidos Biomedical Research, Inc., Frederick National Laboratory for Cancer Research, Frederick, Maryland, United States of America

Pamela A. Kozlowski

Affiliation Department of Microbiology, Immunology and Parasitology, Louisiana State University Health Sciences Center, New Orleans, Louisiana, United States of America

Ronald C. Desrosiers

Affiliation Department of Pathology, University of Miami Miller School of Medicine, Miami, Florida, United States of America

Ashley T. Haase

Affiliation Department of Microbiology, Medical School, University of Minnesota, MMC 196, 420 Delaware Street S.E., Minneapolis, Minnesota, United States of America

R. Paul Johnson

rpaul.johnson@emory.edu

Affiliations Yerkes National Primate Research Center, Emory University, Atlanta GA, United States of America, New England Primate Research Center, Harvard Medical School, Southborough Campus, Pine Hill Drive, Southborough, MA, United States of America, Ragon Institute of MGH, MIT and Harvard, Cambridge, MA, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: SA RKR ATH RPJ. Data curation: SA RKR JG FEW YY YL MP JDL WL PAK ATH RPJ. Formal analysis: SA RKR JG FEW YY QL MC PAK ATH RPJ. Funding acquisition: JDL RCD ATH RPJ. Investigation: SA RKR JG FEW YY JVC QL MC YL MP BFK PAK. Project administration: SA RKR JG MP ATH RPJ. Resources: RCD. Supervision: SA RKR JG JDL PAK ATH RPJ. Visualization: SA RKR PAK. Writing – original draft: SA RKR RPJ. Writing – review & editing: SA RKR JDL PAK RCD ATH RPJ.