About the Authors

Naomi H. Philip

Affiliations University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America

Alexandra DeLaney

Affiliation University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America

Lance W. Peterson

Affiliations University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America

Melanie Santos-Marrero

Affiliations University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America

Jennifer T. Grier

Affiliation University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America

Yan Sun

Affiliation University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America

Meghan A. Wynosky-Dolfi

Affiliation University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America

Erin E. Zwack

Affiliation University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America

Baofeng Hu

Affiliation University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America

Tayla M. Olsen

Affiliation University of Washington, Department of Immunology, Seattle, Washington, United States of America

Anthony Rongvaux

Affiliation Fred Hutchinson Cancer Research Center, Clinical Research Division and Program in Immunology, Seattle, Washington, United States of America

Scott D. Pope

Affiliation Yale University School of Medicine, Department of Immunobiology, New Haven, Connecticut, United States of America

Carolina B. López

Affiliations University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America

Andrew Oberst

Affiliation University of Washington, Department of Immunology, Seattle, Washington, United States of America

Daniel P. Beiting

Affiliations University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America

Jorge Henao-Mejia

Affiliations University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America, Department of Pathology and Laboratory Medicine, Perelman School of Medicine at the University of Pennsylvania and Children’s Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Igor E. Brodsky

ibrodsky@vet.upenn.edu

Affiliations University of Pennsylvania School of Veterinary Medicine, Department of Pathobiology, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania Perelman School of Medicine, Institute for Immunology, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: IEB NHP AO DPB. Data curation: NHP DPB. Formal analysis: IEB NHP DPB AD. Funding acquisition: IEB AO CBL. Investigation: IEB NHP AD LWP MSM JTG YS MAWD EEZ BH. Methodology: NHP DPB JHM SDP AD. Resources: AR TMO AO. Supervision: IEB. Writing – original draft: IEB NHP. Writing – review & editing: IEB NHP.