About the Authors

Alan M. Watson

alan.watson@pitt.edu

Affiliation Center for Vaccine Research, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

L. K. Metthew Lam

Affiliation Center for Vaccine Research, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

William B. Klimstra

Affiliation Center for Vaccine Research, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Kate D. Ryman

† Deceased.

Affiliation Center for Vaccine Research, Department of Microbiology and Molecular Genetics, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AMW LKML WBK KDR. Performed the experiments: AMW LKML. Analyzed the data: AMW LKML KDR. Wrote the paper: AMW WBK KDR.