About the Authors

Philana Ling Lin

philana.lin@chp.edu (PLL); joanne@pitt.edu (JLF)

Affiliation Department of Pediatrics, Children’s Hospital of Pittsburgh of University of Pittsburgh Medical Center, Pittsburgh, Pennsylvania, United States of America

Pauline Maiello

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Hannah P. Gideon

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

M. Teresa Coleman

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Anthony M. Cadena

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Mark A. Rodgers

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Robert Gregg

Affiliations Department of Pediatrics, Children’s Hospital of Pittsburgh of University of Pittsburgh Medical Center, Pittsburgh, Pennsylvania, United States of America, Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Melanie O’Malley

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Jaime Tomko

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Daniel Fillmore

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

L. James Frye

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Tara Rutledge

Affiliation Department of Pediatrics, Children’s Hospital of Pittsburgh of University of Pittsburgh Medical Center, Pittsburgh, Pennsylvania, United States of America

Robert M. DiFazio

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Christopher Janssen

Affiliation Division of Laboratory Animal Resources, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Edwin Klein

Affiliation Division of Laboratory Animal Resources, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Peter L. Andersen

Affiliation Department of Infectious Diseases Immunology, Statens Serum Institute, Copenhagen, Denmark

Sarah M. Fortune

Affiliation Department of Immunology and Infectious Diseases, Harvard School of Public Health, Boston, Massachusetts, United States of America

JoAnne L. Flynn

philana.lin@chp.edu (PLL); joanne@pitt.edu (JLF)

Affiliation Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, Pennsylvania, United States of America

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: PLA is the co-inventor of the H56 vaccine patent, but all rights have been assigned to the Serum Staten Institute.

Author Contributions

Conceived and designed the experiments: PLL JLF SMF. Performed the experiments: HPG AMC MAR JT DF LJF MO TR RMD CJ EK PLL. Analyzed the data: PM HPG MTC AMC MAR RG TR RMD PLA SMF PLL JLF. Contributed reagents/materials/analysis tools: PM HPG MTC AMC RG SMF PLA. Wrote the paper: PLL JLF SMF PM.