About the Authors

Christopher M. Ziegler

Affiliations Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America, Cellular, Molecular, and Biomedical Sciences Graduate Program, University of Vermont, Burlington, Vermont, United States of America

Philip Eisenhauer

Affiliation Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America

Emily A. Bruce

Affiliation Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America

Marion E. Weir

Affiliation Department of Biology, University of Vermont, Burlington, Vermont, United States of America

Benjamin R. King

Affiliations Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America, Cellular, Molecular, and Biomedical Sciences Graduate Program, University of Vermont, Burlington, Vermont, United States of America

Joseph P. Klaus

Current address: Department of Immunology and Microbial Science, The Scripps Research Institute, La Jolla, California, United States of America

Affiliations Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America, Cellular, Molecular, and Biomedical Sciences Graduate Program, University of Vermont, Burlington, Vermont, United States of America

Dimitry N. Krementsov

Affiliation Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America

David J. Shirley

Affiliation Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America

Bryan A. Ballif

Affiliation Department of Biology, University of Vermont, Burlington, Vermont, United States of America

Jason Botten

jbotten@uvm.edu

Affiliations Department of Medicine, Division of Immunobiology, University of Vermont, Burlington, Vermont, United States of America, Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CMZ EAB BAB JPK PE JB. Performed the experiments: CMZ PE MEW BRK JPK BAB JB. Analyzed the data: CMZ EAB MEW BRK JPK DJS BAB JB. Contributed reagents/materials/analysis tools: DJS DNK BAB JB. Wrote the paper: CMZ EAB BAB JB. Edited the manuscript: CMZ PE EAB MEW BRK JPK DJS BAB JB. Oversaw all aspects of this research project: JB.