About the Authors

Fang Lu

Affiliation The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Horng-Shen Chen

Affiliation The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Andrew V. Kossenkov

Affiliation The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Karen DeWispeleare

Affiliation The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Kyoung-Jae Won

wonk@mail.med.upenn.edu (KJW); Lieberman@wistar.org (PML)

Affiliation The Institute for Diabetes Obesity and Metabolism, Department of Genetics, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Paul M. Lieberman

wonk@mail.med.upenn.edu (KJW); Lieberman@wistar.org (PML)

Affiliation The Wistar Institute, Philadelphia, Pennsylvania, United States of America

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: PML is a founder of a company with interest in EBV associated disease. This does not alter our adherence to all PLOS policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: FL KJW PML. Performed the experiments: FL KD HSC. Analyzed the data: FL AVK KJW PML. Contributed reagents/materials/analysis tools: KJW AVK PML. Wrote the paper: FL KJW AVK PML.