About the Authors

Elise Courtot

Affiliations INRA, UMR1282 Infectiologie et Santé Publique, Nouzilly, France, Université de François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Claude L. Charvet

Affiliations INRA, UMR1282 Infectiologie et Santé Publique, Nouzilly, France, Université de François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Robin N. Beech

Affiliations INRA, UMR1282 Infectiologie et Santé Publique, Nouzilly, France, Institute of Parasitology, McGill University, Macdonald Campus, Sainte Anne de Bellevue, Québec, Canada

Abdallah Harmache

Affiliations INRA, UMR1282 Infectiologie et Santé Publique, Nouzilly, France, Université de François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Adrian J. Wolstenholme

Affiliation Department of Infectious Disease and Center for Tropical and Emerging Global Disease, University of Georgia, Athens, Georgia, United States of America

Lindy Holden-Dye

Affiliation School of Biological Sciences, University of Southampton, Southampton, United Kingdom

Vincent O’Connor

Affiliation School of Biological Sciences, University of Southampton, Southampton, United Kingdom

Nicolas Peineau

Affiliation Université François Rabelais de Tours, Département de physiologie animale, Tours, France

Debra J. Woods

Affiliation Veterinary Medicine Research and Development, Zoetis LLC, Kalamazoo, Michigan, United States of America

Cedric Neveu

cedric.neveu@tours.inra.fr

Affiliations INRA, UMR1282 Infectiologie et Santé Publique, Nouzilly, France, Université de François Rabelais de Tours, UMR1282 Infectiologie et Santé Publique, Tours, France

Competing Interests

DJW is employed by Zoetis, which is developing veterinary pharmaceuticals including anthelmintics. The present study was performed as a collaborative project between the French National Institute for Agronomical Research and Zoetis. Accordingly, the academic institution received a specific research grant from Zoetis. This does not alter our adherence to all PLOS policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: EC CLC AH RNB LHD VO DJW CN. Performed the experiments: EC CLC AH RNB CN. Analyzed the data: EC CLC AH RNB LHD VO AJW NP CN. Contributed reagents/materials/analysis tools: CLC AH NP LHD VO AJW. Wrote the paper: EC CLC RNB LHD VO AJW CN.