About the Authors

James M. Billingsley

Affiliations Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Premeela A. Rajakumar

Affiliations Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Michelle A. Connole

Affiliation Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America

Nadine C. Salisch

Affiliations Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America, Crucell Holland BV, Leiden, The Netherlands

Sama Adnan

Affiliations Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia, United States of America

Yury V. Kuzmichev

Affiliation Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America

Henoch S. Hong

Affiliation Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America

R. Keith Reeves

Affiliations Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America, Center for Virology and Vaccine Research, Beth Israel Deaconess Medical Center, Boston, Massachusetts, United States of America

Hyung-joo Kang

Affiliation Division of Preventive and Behavioral Medicine, University of Massachusetts Medical Center, Worcester, Massachusetts, United States of America

Wenjun Li

Affiliation Division of Preventive and Behavioral Medicine, University of Massachusetts Medical Center, Worcester, Massachusetts, United States of America

Qingsheng Li

Affiliation Nebraska Center for Virology and School of Biological Sciences, University of Nebraska-Lincoln, Lincoln, Nebraska, United States of America

Ashley T. Haase

Affiliation University of Minnesota, Microbiology Department, Minneapolis, Minnesota, United States of America

R. Paul Johnson

rpaul.johnson@emory.edu

Affiliations Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough, Massachusetts, United States of America, Yerkes National Primate Research Center, Emory University, Atlanta, Georgia, United States of America, Ragon Institute of MGH, MIT, and Harvard, Cambridge, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RPJ JMB PAR NCS YVK HSH. Performed the experiments: JMB PAR MAC. Analyzed the data: RPJ JMB PAR MAC HjK WL SA. Contributed reagents/materials/analysis tools: RKR QL ATH. Wrote the paper: RPJ JMB PAR MAC.