About the Authors

Haiying Li

Affiliation Center for Virology and Vaccine Research, Beth Israel Deaconess Medical Center, Boston, Massachusetts, United States of America

Laura E. Richert-Spuhler

Affiliation Department of Pharmaceutics, Washington National Primate Research Center, University of Washington, Seattle, Washington, United States of America

Tristan I. Evans

Affiliation Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough Campus, Southborough, Massachusetts, United States of America

Jacqueline Gillis

Affiliation Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough Campus, Southborough, Massachusetts, United States of America

Michelle Connole

Affiliation Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough Campus, Southborough, Massachusetts, United States of America

Jacob D. Estes

Affiliation AIDS and Cancer Virus Program, Leidos Biomedical Research, Inc., Frederick National Laboratory, Frederick, Maryland, United States of America

Brandon F. Keele

Affiliation AIDS and Cancer Virus Program, Leidos Biomedical Research, Inc., Frederick National Laboratory, Frederick, Maryland, United States of America

Nichole R. Klatt

Affiliation Department of Pharmaceutics, Washington National Primate Research Center, University of Washington, Seattle, Washington, United States of America

R. Keith Reeves

rreeves@bidmc.harvard.edu

Affiliations Center for Virology and Vaccine Research, Beth Israel Deaconess Medical Center, Boston, Massachusetts, United States of America, Division of Immunology, New England Primate Research Center, Harvard Medical School, Southborough Campus, Southborough, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RKR. Performed the experiments: HL TIE JG MC LER. Analyzed the data: HL NRK RKR. Contributed reagents/materials/analysis tools: JDE BFK NRK. Wrote the paper: HL RKR.