About the Authors

Yi-Pin Lin

Affiliation Department of Molecular Biology and Microbiology, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Vivian Benoit

Current address: Program in Molecular Medicine, University of Massachusetts Medical School, Worcester, Massachusetts, United States of America

Affiliation Department of Molecular Biology and Microbiology, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Xiuli Yang

Affiliations Department of Veterinary Medicine, University of Maryland, College Park, Maryland, United States of America, Virginia–Maryland Regional College of Veterinary Medicine, College Park, Maryland, United States of America

Raúl Martínez-Herranz

Affiliation Department of Molecular Biology and Microbiology, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Utpal Pal

Affiliations Department of Veterinary Medicine, University of Maryland, College Park, Maryland, United States of America, Virginia–Maryland Regional College of Veterinary Medicine, College Park, Maryland, United States of America

John M. Leong

John.Leong@tufts.edu

Affiliation Department of Molecular Biology and Microbiology, Tufts University School of Medicine, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YPL VB XY UP JML. Performed the experiments: YPL VB XY RMH. Analyzed the data: YPL XY UP JML. Contributed reagents/materials/analysis tools: UP JML. Wrote the paper: YPL XY UP JML.