About the Authors

Sandra J. Raffel

Affiliation Laboratory of Zoonotic Pathogens, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Disease, National Institutes of Health, Hamilton, Montana, United States of America

James M. Battisti

Affiliations Laboratory of Zoonotic Pathogens, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Disease, National Institutes of Health, Hamilton, Montana, United States of America, Division of Biological Sciences, The University of Montana, Missoula, Montana, United States of America

Robert J. Fischer

Affiliation Laboratory of Zoonotic Pathogens, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Disease, National Institutes of Health, Hamilton, Montana, United States of America

Tom G. Schwan

tschwan@niaid.nih.gov

Affiliation Laboratory of Zoonotic Pathogens, Rocky Mountain Laboratories, National Institute of Allergy and Infectious Disease, National Institutes of Health, Hamilton, Montana, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJR JMB RJF TGS. Performed the experiments: SJR JMB RJF TGS. Analyzed the data: SJR TGS. Contributed reagents/materials/analysis tools: TGS. Wrote the paper: SJR TGS.