About the Authors

Christopher D. Dupont

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

David A. Christian

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Elizabeth M. Selleck

Affiliation Department of Molecular Microbiology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Marion Pepper

Affiliation Department of Immunology, University of Washington, Seattle, Washington, United States of America

Michael Leney-Greene

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gretchen Harms Pritchard

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Anita A. Koshy

Affiliation Department of Neurology, University of Arizona, Tucson, Arizona, United States of America

Sagie Wagage

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Morgan A. Reuter

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

L. David Sibley

Affiliation Department of Molecular Microbiology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Michael R. Betts

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Christopher A. Hunter

chunter@vet.upenn.edu

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CDD DAC CAH EMS LDS. Performed the experiments: CDD EMS DAC MAR GHP MLG SW. Analyzed the data: CDD EMS MLG GHP DAC MAR. Contributed reagents/materials/analysis tools: MRB AAK MP LDS. Wrote the paper: CDD CAH EMS.