About the Authors

Scott M. Robinson

Contributed equally to this work with: Scott M. Robinson, Ginger Tsueng, Jon Sin

SMR, GT, and JS are joint first authors on this work.

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Ginger Tsueng

Contributed equally to this work with: Scott M. Robinson, Ginger Tsueng, Jon Sin

SMR, GT, and JS are joint first authors on this work.

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Jon Sin

Contributed equally to this work with: Scott M. Robinson, Ginger Tsueng, Jon Sin

SMR, GT, and JS are joint first authors on this work.

Affiliation Donald P. Shiley BioScience Center, San Diego State University, San Diego, California, United States of America

Vrushali Mangale

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Shahad Rahawi

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Laura L. McIntyre

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Wesley Williams

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Nelson Kha

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Casey Cruz

Affiliation Department of Biology and Center for Microbial Sciences, San Diego State University, San Diego, California, United States of America

Bryan M. Hancock

Affiliation Department of Biology and Center for Microbial Sciences, San Diego State University, San Diego, California, United States of America

David P. Nguyen

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

M. Richard Sayen

Affiliation Donald P. Shiley BioScience Center, San Diego State University, San Diego, California, United States of America

Brett J. Hilton

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Kelly S. Doran

Affiliation Department of Biology and Center for Microbial Sciences, San Diego State University, San Diego, California, United States of America

Anca M. Segall

Affiliation Department of Biology and Center for Microbial Sciences, San Diego State University, San Diego, California, United States of America

Roland Wolkowicz

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Christopher T. Cornell

Current address: Global Medical Affairs, Allergan, Inc., Irvine, California, United States of America.

Affiliation Department of Immunology and Microbial Science, SP30-2110, The Scripps Research Institute, La Jolla, California, United States of America

J. Lindsay Whitton

Affiliation Department of Immunology and Microbial Science, SP30-2110, The Scripps Research Institute, La Jolla, California, United States of America

Roberta A. Gottlieb

Affiliation Donald P. Shiley BioScience Center, San Diego State University, San Diego, California, United States of America

Ralph Feuer

rfeuer@sciences.sdsu.edu

Affiliation The Integrated Regenerative Research Institute (IRRI) at San Diego State University, Cell & Molecular Biology Joint Doctoral Program, Department of Biology, San Diego State University, San Diego, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SMR GT JS RAG RF. Performed the experiments: SMR GT JS VM SR LLM WW NK CC BMH DPN. Analyzed the data: SMR GT JS JLW RAG RF. Contributed reagents/materials/analysis tools: MRS BJH KSD AMS RW CTC JLW RAG. Wrote the paper: SMR GT JS RAG RF.