About the Authors

Christopher Paul Walczak

Affiliations Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America, Cellular and Molecular Biology Graduate Program, Ann Arbor, Michigan, United States of America

Madhu Sudhan Ravindran

Affiliation Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Takamasa Inoue

Affiliation Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Billy Tsai

btsai@umich.edu

Affiliation Department of Cell and Developmental Biology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CPW MSR BT. Performed the experiments: CPW MSR TI. Analyzed the data: CPW MSR TI BT. Contributed reagents/materials/analysis tools: CPW MSR TI BT. Wrote the paper: CPW MSR TI BT.