About the Authors

Asher Maroof

Current address: UCB Pharma Ltd, Slough, United Kingdom.

Affiliation GlaxoSmithKline Vaccines, Hamilton, Montana, United States of America

Yvonne M. Yorgensen

Affiliation GlaxoSmithKline Vaccines, Hamilton, Montana, United States of America

Yufeng Li

Affiliation GlaxoSmithKline Vaccines, Hamilton, Montana, United States of America

Jay T. Evans

jay.t.evans@gsk.com

Affiliation GlaxoSmithKline Vaccines, Hamilton, Montana, United States of America

Competing Interests

All authors were employed by the GlaxoSmithKline group of companies GSK Vaccines during completion of the experimental studies contained herein. This does not alter our adherence to all PLoS Pathogens policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JTE YMY YL AM. Performed the experiments: YMY YL AM. Analyzed the data: JTE YMY YL AM. Contributed reagents/materials/analysis tools: JTE YMY YL AM. Wrote the paper: JTE YMY YL AM.