About the Authors

Kei Sato

ksato@virus.kyoto-u.ac.jp (KS); ykoyanag@virus.kyoto-u.ac.jp (YK)

Affiliations Laboratory of Virus Pathogenesis, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan, Center for Emerging Virus Research, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan

Naoko Misawa

Affiliation Laboratory of Virus Pathogenesis, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan

Shingo Iwami

Affiliations Department of Biology, Faculty of Sciences, Kyushu University, Fukuoka, Fukuoka, Japan, Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency (JST), Kawaguchi, Saitama, Japan

Yorifumi Satou

Affiliation Laboratory of Viral Control, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan

Masao Matsuoka

Affiliation Laboratory of Viral Control, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan

Yukihito Ishizaka

Affiliation Department of Intractable Diseases, National Center for Global Health and Medicine, Shinjuku-ku, Tokyo, Japan

Mamoru Ito

Affiliation Central Institute for Experimental Animals, Kawasaki, Kanagawa, Japan

Kazuyuki Aihara

Affiliations Institute of Industrial Science, The University of Tokyo, Meguro-ku, Tokyo, Japan, Graduate School of Information Science and Technology, The University of Tokyo, Meguro-ku, Tokyo, Japan

Dong Sung An

Affiliations Division of Hematology-Oncology, University of California, Los Angeles (UCLA), Los Angeles, California, United States of America, School of Nursing, UCLA, Los Angeles, California, United States of America, AIDS Institute, UCLA, Los Angeles, California, United States of America

Yoshio Koyanagi

ksato@virus.kyoto-u.ac.jp (KS); ykoyanag@virus.kyoto-u.ac.jp (YK)

Affiliations Laboratory of Virus Pathogenesis, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan, Center for Emerging Virus Research, Institute for Virus Research, Kyoto University, Kyoto, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KS YK. Performed the experiments: KS NM YS. Analyzed the data: KS NM SI YS MM. Contributed reagents/materials/analysis tools: SI MM YI MI KA DSA. Wrote the paper: KS YK.