About the Authors

Michael J. McClellan

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

C. David Wood

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

Opeoluwa Ojeniyi

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

Tim J. Cooper

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

Aditi Kanhere

Current address: School of Biosciences, University of Birmingham, Edgbaston, Birmingham, United Kingdom.

Affiliation MRC Centre for Medical Molecular Virology, Division of Infection and Immunity, Paul O'Gorman Building, University College London, London, United Kingdom

Aaron Arvey

Affiliation Memorial Sloan-Kettering Cancer Center, New York, New York, United States of America

Helen M. Webb

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

Richard D. Palermo

Current address: London Research Institute, Lincoln's Inn Fields Laboratories, London, United Kingdom.

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

Marie L. Harth-Hertle

Affiliation Department of Gene Vectors, Helmholtz Center Munich, German Research Center for Environmental Health, Munich, Germany

Bettina Kempkes

Affiliation Department of Gene Vectors, Helmholtz Center Munich, German Research Center for Environmental Health, Munich, Germany

Richard G. Jenner

Affiliation MRC Centre for Medical Molecular Virology, Division of Infection and Immunity, Paul O'Gorman Building, University College London, London, United Kingdom

Michelle J. West

M.J.West@sussex.ac.uk

Affiliation School of Life Sciences, John Maynard-Smith Building, University of Sussex, Falmer, Brighton, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MJW MJM CDW. Performed the experiments: MJM CDW OO TJC HMW RDP. Analyzed the data: AK RGJ AA MJM MJW. Contributed reagents/materials/analysis tools: MLHH BK. Wrote the paper: MJW RGJ.