About the Authors

Kimberly M. Brothers

Current address: University of Pittsburgh Medical Center, Department of Ophthamology, Eye and Ear Institute, Pittsburgh, Pennsylvania, United States of America.

Affiliations Department of Molecular and Biomedical Sciences, University of Maine, Orono, Maine, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine, United States of America

Remi L. Gratacap

Affiliation Department of Molecular and Biomedical Sciences, University of Maine, Orono, Maine, United States of America

Sarah E. Barker

Affiliation Department of Molecular and Biomedical Sciences, University of Maine, Orono, Maine, United States of America

Zachary R. Newman

Affiliation Department of Molecular and Biomedical Sciences, University of Maine, Orono, Maine, United States of America

Ashley Norum

Current address: Pacific University, Forest Grove, Oregon, United States of America.

Affiliation Department of Molecular and Biomedical Sciences, University of Maine, Orono, Maine, United States of America

Robert T. Wheeler

robert.wheeler@umit.maine.edu

Affiliations Department of Molecular and Biomedical Sciences, University of Maine, Orono, Maine, United States of America, Graduate School of Biomedical Sciences and Engineering, University of Maine, Orono, Maine, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KMB RLG SEB RTW. Performed the experiments: KMB RLG SEB ZRN AN. Analyzed the data: KMB RLG SEB ZRN AN. Wrote the paper: KMB RTW.