About the Authors

Fei Liu

Affiliation Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America

Matthew Campagna

Affiliations Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America, Institute for Hepatitis and Virology Research, Hepatitis B Foundation, Doylestown, Pennsylvania, United States of America

Yonghe Qi

Affiliation National Institute of Biological Sciences, Beijing, People's Republic of China

Xuesen Zhao

Affiliation Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America

Fang Guo

Affiliation Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America

Chunxiao Xu

Affiliation Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America

Sichen Li

Affiliation Neurology Department, David Geffen Medical School, University of Los Angeles, Los Angeles, California, United States of America

Wenhui Li

Affiliation National Institute of Biological Sciences, Beijing, People's Republic of China

Timothy M. Block

Affiliations Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America, Institute for Hepatitis and Virology Research, Hepatitis B Foundation, Doylestown, Pennsylvania, United States of America

Jinhong Chang

Affiliation Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America

Ju-Tao Guo

Ju-Tao.Guo@drexelmed.edu

Affiliation Drexel Institute for Biotechnology and Virology Research, Department of Microbiology and Immunology, Drexel University College of Medicine, Doylestown, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL JTG. Performed the experiments: FL MC YQ XZ FG CX JC. Analyzed the data: FL YQ SL WL TMB JC JTG. Wrote the paper: FL JC JTG.