About the Authors

Stephanie Traub

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Alexandra Nikonova

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Alan Carruthers

Affiliation MedImmune Ltd, Cambridge, United Kingdom

Rebecca Dunmore

Affiliation MedImmune Ltd, Cambridge, United Kingdom

Katherine A. Vousden

Affiliation MedImmune Ltd, Cambridge, United Kingdom

Leila Gogsadze

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Weidong Hao

Affiliation MedImmune LLC, Gaithersburg, Maryland, United States of America

Qing Zhu

Affiliation MedImmune LLC, Gaithersburg, Maryland, United States of America

Katie Bernard

Affiliation MedImmune LLC, Gaithersburg, Maryland, United States of America

Jie Zhu

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Michael Dymond

Affiliation AstraZeneca, Charnwood, United Kingdom

Gary R. McLean

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Ross P. Walton

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Nicholas Glanville

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Alison Humbles

Affiliation MedImmune LLC, Gaithersburg, Maryland, United States of America

Musa Khaitov

Affiliation NRC Institute of Immunology FMBA, Moscow, Russia

Ted Wells

Affiliation AstraZeneca, Charnwood, United Kingdom

Roland Kolbeck

Affiliation MedImmune LLC, Gaithersburg, Maryland, United States of America

Andrew J. Leishman

Affiliation AstraZeneca, Charnwood, United Kingdom

Matthew A. Sleeman

Affiliation MedImmune Ltd, Cambridge, United Kingdom

Nathan W. Bartlett

Contributed equally to this work with: Nathan W. Bartlett, Sebastian L. Johnston

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Sebastian L. Johnston

Contributed equally to this work with: Nathan W. Bartlett, Sebastian L. Johnston

s.johnston@imperial.ac.uk

Affiliation National Heart and Lung Institute, MRC and Asthma UK Centre in Allergic Mechanisms of Asthma, Centre for Respiratory Infection, Imperial College London, London, United Kingdom

Competing Interests

The following authors (AC, RD, KAV, WH, QZ, AH, RK, MAS, KB, MD, TW, AJL) are full time employees of MedImmune Ltd. or MedImmune LLC, wholly owned subsidiaries of Astrazeneca PLC. As employees they also receive shares and share options in the Astrazeneca Group. During the course of the study the following authors (AJL, TW) were employees of Astrazeneca PLC and received shares and share options. Astrazeneca is a global pharmaceutical company that conducts basic research and clinical trials in respiratory diseases such as severe asthma and chronic obstructive pulmonary disease. This work was in part funded by MedImmune and Astrazeneca PLC. This does not alter our adherence to all PLoS Pathogens policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SLJ NWB ST MAS AC RK AJL AH TW. Performed the experiments: ST NWB RPW NG AN LG JZ GRM AC RK AH RD QZ WH KB MD. Analyzed the data: ST NWB SLJ AC RD RK MAS AJL AH. Contributed reagents/materials/analysis tools: MAS KAV MK. Wrote the paper: ST NWB SLJ MAS KAV.