About the Authors

James P. Hewitson

Affiliation Institute of Immunology and Infection Research, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Al C. Ivens

Affiliation Centre for Immunity, Infection and Evolution, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Yvonne Harcus

Affiliation Institute of Immunology and Infection Research, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Kara J. Filbey

Affiliation Institute of Immunology and Infection Research, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Henry J. McSorley

Affiliation Institute of Immunology and Infection Research, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Janice Murray

Affiliation Institute of Immunology and Infection Research, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Stephen Bridgett

Affiliation Gene Pool, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

David Ashford

Affiliation Technology Facility, University of York, York, United Kingdom

Adam A. Dowle

Affiliation Technology Facility, University of York, York, United Kingdom

Rick M. Maizels

rick.maizels@ed.ac.uk

Affiliation Institute of Immunology and Infection Research, Ashworth Laboratories, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JPH RMM. Performed the experiments: JPH YH KJF HJM JM AAD. Analyzed the data: JPH ACI SB DA AAD RMM. Wrote the paper: JPH RMM.