About the Authors

Stephen G. Preston

Affiliations Nuffield Department of Surgical Sciences, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom, Department of Zoology, University of Oxford, Oxford, United Kingdom

Juraj Majtán

Affiliation Institute of Zoology, Slovak Academy of Sciences, Bratislava, Slovakia

Chrisoula Kouremenou

Affiliation Nuffield Department of Surgical Sciences, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Oliwia Rysnik

Affiliation Nuffield Department of Clinical Medicine, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Lena F. Burger

Affiliation Nuffield Department of Surgical Sciences, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Alejandro Cabezas Cruz

Affiliation University of South Bohemia, Faculty of Science and Biology Centre of the ASCR, Institute of Parisitology, České Budějovice, Czech Republic

Maylin Chiong Guzman

Affiliation Center for Genetic Engineering and Biotechnology, Animal Biotechnology Division, Havana, Cuba

Miles A. Nunn

Affiliation NERC Centre for Ecology & Hydrology, Crowmarsh Gifford, Wallingford, Oxfordshire, United Kingdom

Guido C. Paesen

Affiliation Division of Structural Biology, Henry Wellcome Building for Genomic Medicine, Oxford, United Kingdom

Patricia A. Nuttall

Affiliations Department of Zoology, University of Oxford, Oxford, United Kingdom, NERC Centre for Ecology & Hydrology, Crowmarsh Gifford, Wallingford, Oxfordshire, United Kingdom

Jonathan M. Austyn

jon.austyn@nds.ox.ac.uk

Affiliation Nuffield Department of Surgical Sciences, University of Oxford, John Radcliffe Hospital, Oxford, United Kingdom

Competing Interests

I have read the journal's policy and have the following conflicts. Some of the data in this publication have been used in the patent applications WO2010/032008 and WO2011/117582 which are owned by the Natural Environment Research Council (NERC) in agreement with the University of Oxford, and were licenced to IXO Therapeutics Ltd. until 31st Dec 2012; the licence has now expired and there are currently no commercial activities relating to these patents. SGP, CK, PAN and JMA were formerly shareholders of, and SGP, GCP and JMA were consultants to, IXO Therapeutics Ltd.; this spin-out company is no longer operational. This does not alter our adherence to all PLoS Pathogens policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SGP MAN GCP PAN JMA. Performed the experiments: SGP JM CK OR LFB ACC MCG GCP. Analyzed the data: SGP JM CK OR LFB ACC MCG GCP PAN JMA. Contributed reagents/materials/analysis tools: SGP CK JM LFB ACC MCG GCP. Wrote the paper: SGP PAN JMA.