About the Authors

Marion Avril

Affiliation Seattle Biomedical Research Institute, Seattle, Washington, United States of America

Andrew J. Brazier

Affiliation Seattle Biomedical Research Institute, Seattle, Washington, United States of America

Martin Melcher

Affiliation Seattle Biomedical Research Institute, Seattle, Washington, United States of America

Sowmya Sampath

Affiliation Seattle Biomedical Research Institute, Seattle, Washington, United States of America

Joseph D. Smith

joe.smith@seattlebiomed.org

Affiliations Seattle Biomedical Research Institute, Seattle, Washington, United States of America, Department of Global Health, University of Washington, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MA AJB MM JDS. Performed the experiments: MA AJB MM SS. Analyzed the data: MA AJB MM JDS SS. Contributed reagents/materials/analysis tools: MA AJB MM. Wrote the paper: MA JDS.