About the Authors

Anat Melamed

Affiliation Department of Immunology, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Daniel J. Laydon

Affiliation Department of Immunology, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Nicolas A. Gillet

Affiliations Department of Immunology, Wright-Fleming Institute, Imperial College London, London, United Kingdom, Molecular and Cellular Epigenetics, Interdisciplinary Cluster for Applied Genoproteomics (GIGA) of University of Liège (ULg), Liège, Belgium

Yuetsu Tanaka

Affiliation Graduate School and Faculty of Medicine, University of the Ryukyus, Okinawa, Japan

Graham P. Taylor

Affiliation Department of Genitourinary Medicine and Communicable Diseases, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Charles R. M. Bangham

c.bangham@imperial.ac.uk

Affiliation Department of Immunology, Wright-Fleming Institute, Imperial College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AM GPT CRMB. Performed the experiments: AM NAG. Analyzed the data: AM. Contributed reagents/materials/analysis tools: YT GPT NAG DJL. Wrote the paper: AM CRMB. Clinical diagnosis and assessment: GPT. Designed the statistical analysis: AM DJL CRMB.