About the Authors

Travis J. Wiles

Affiliation Division of Microbiology and Immunology, Pathology Department, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

J. Paul Norton

Contributed equally to this work with: J. Paul Norton, Sara N. Smith

Affiliation Division of Microbiology and Immunology, Pathology Department, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Sara N. Smith

Contributed equally to this work with: J. Paul Norton, Sara N. Smith

Affiliation Department of Microbiology and Immunology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Adam J. Lewis

Affiliation Division of Microbiology and Immunology, Pathology Department, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Harry L. T. Mobley

Affiliation Department of Microbiology and Immunology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Sherwood R. Casjens

Affiliation Division of Microbiology and Immunology, Pathology Department, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Matthew A. Mulvey

mulvey@path.utah.edu

Affiliation Division of Microbiology and Immunology, Pathology Department, University of Utah School of Medicine, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TJW SRC MAM. Performed the experiments: TJW JPN SNS SRC AJL. Analyzed the data: TJW SRC MAM. Contributed reagents/materials/analysis tools: TJW JPN SNS HLTM SRC MAM. Wrote the paper: TJW JPN SRC MAM.