About the Authors

Cheng-Der Liu

Contributed equally to this work with: Cheng-Der Liu, Ya-Lin Chen

Affiliation Department of Life Sciences, Tzu-Chi University, Hualien, Taiwan

Ya-Lin Chen

Contributed equally to this work with: Cheng-Der Liu, Ya-Lin Chen

Affiliation Department of Life Sciences, Tzu-Chi University, Hualien, Taiwan

Yi-Li Min

Affiliation Department of Life Sciences, Tzu-Chi University, Hualien, Taiwan

Bo Zhao

Affiliation Department of Medicine and Microbiology and Molecular Genetics, Channing Laboratory, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America

Chi-Ping Cheng

Affiliation Department of Life Sciences, Tzu-Chi University, Hualien, Taiwan

Myung-Soo Kang

Affiliation Department of Medicine and Microbiology and Molecular Genetics, Channing Laboratory, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America

Shu-Jun Chiu

Affiliation Department of Life Sciences, Tzu-Chi University, Hualien, Taiwan

Elliott Kieff

Affiliation Department of Medicine and Microbiology and Molecular Genetics, Channing Laboratory, Brigham and Women's Hospital and Harvard Medical School, Boston, Massachusetts, United States of America

Chih-Wen Peng

pengcw@mail.tcu.edu.tw

Affiliation Department of Life Sciences, Tzu-Chi University, Hualien, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CWP. Performed the experiments: CDL YLC YLM BZ MSK. Analyzed the data: CPC SJC. Contributed reagents/materials/analysis tools: EK. Wrote the paper: CWP BZ.