About the Authors

Matthew R. Olson

Contributed equally to this work with: Matthew R. Olson, Daniel S. McDermott

Current address: Department of Microbiology and Immunology, University of Melbourne, Parkville, Victoria, Australia

Affiliation Department of Microbiology, University of Iowa, Iowa City, Iowa, United States of America

Daniel S. McDermott

Contributed equally to this work with: Matthew R. Olson, Daniel S. McDermott

Affiliation Interdisciplinary Graduate Program in Immunology, University of Iowa, Iowa City, Iowa, United States of America

Steven M. Varga

steven-varga@uiowa.edu

Affiliations Department of Microbiology, University of Iowa, Iowa City, Iowa, United States of America, Interdisciplinary Graduate Program in Immunology, University of Iowa, Iowa City, Iowa, United States of America, Department of Pathology, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MRO DSM SMV. Performed the experiments: MRO DSM SMV. Analyzed the data: MRO DSM SMV. Contributed reagents/materials/analysis tools: MRO DSM SMV. Wrote the paper: MRO DSM SMV.