About the Authors

Hongguang Shao

Contributed equally to this work with: Hongguang Shao, Xinshe Li

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Xinshe Li

Contributed equally to this work with: Hongguang Shao, Xinshe Li

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Thomas J. Nolan

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Holman C. Massey Jr.

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Edward J. Pearce

Affiliation Department of Pathology and Immunology, School of Medicine, Washington University, St. Louis, Missouri, United States of America

James B. Lok

jlok@vet.upenn.edu

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HS XL TJN HCM EJP JBL. Performed the experiments: HS XL TJN. Analyzed the data: HS XL TJN HCM EJP JBL. Contributed reagents/materials/analysis tools: HS XL TJN HCM. Wrote the paper: HS XL TJN HCM EJP JBL.