About the Authors

Pablo R. Murcia

Affiliations Cambridge Infectious Diseases Consortium, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom, Medical Research Council-University of Glasgow Centre for Virus Research, Institute of Infection, Inflammation and Immunity, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Joseph Hughes

Affiliation Medical Research Council-University of Glasgow Centre for Virus Research, Institute of Infection, Inflammation and Immunity, College of Medical, Veterinary and Life Sciences, University of Glasgow, Glasgow, United Kingdom

Patrizia Battista

Affiliation Cambridge Infectious Diseases Consortium, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Lucy Lloyd

Affiliation Animal Health Trust, Centre for Preventive Medicine, Lanwades Park, Newmarket, United Kingdom

Gregory J. Baillie

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Ricardo H. Ramirez-Gonzalez

Affiliation The Genome Analysis Centre, Norwich Research Park, Norwich, United Kingdom

Doug Ormond

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Karen Oliver

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Debra Elton

Affiliation Animal Health Trust, Centre for Preventive Medicine, Lanwades Park, Newmarket, United Kingdom

Jennifer A. Mumford

Affiliation Cambridge Infectious Diseases Consortium, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Mario Caccamo

Affiliation The Genome Analysis Centre, Norwich Research Park, Norwich, United Kingdom

Paul Kellam

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Bryan T. Grenfell

Affiliations Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America, Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, Pennsylvania, United States of America

Edward C. Holmes

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, Pennsylvania, United States of America, Fogarty International Center, National Institute of Health, Bethesda, Maryland, United States of America

James L. N. Wood

jlnw2@cam.ac.uk

Affiliation Cambridge Infectious Diseases Consortium, Department of Veterinary Medicine, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PRM JAM BTG ECH JLNW. Performed the experiments: PRM PB DO KO JH. Analyzed the data: PRM JH ECH. Contributed reagents/materials/analysis tools: DE PK GJB LL MC RHR-G. Wrote the paper: PRM ECH.