About the Authors

Matthew A. Fischer

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Michael L. Davies

Contributed equally to this work with: Michael L. Davies, Irene E. Reider

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Irene E. Reider

Contributed equally to this work with: Michael L. Davies, Irene E. Reider

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Erica L. Heipertz

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Melanie R. Epler

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Janet J. Sei

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Molly A. Ingersoll

Affiliation Department of Gene and Cell Medicine, Mount Sinai School of Medicine, New York, New York, United States of America

Nico Van Rooijen

Affiliation Department of Molecular Cell Biology, Faculty of Medicine, Vrije Universiteit, Amsterdam, The Netherlands

Gwendalyn J. Randolph

Affiliation Department of Gene and Cell Medicine, Mount Sinai School of Medicine, New York, New York, United States of America

Christopher C. Norbury

ccn1@psu.edu

Affiliation Department of Microbiology and Immunology, Pennsylvania State University Milton S. Hershey Medical Center, Hershey, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MAF MLD IER MAI GJR CCN. Performed the experiments: MAF MLD IER CCN. Analyzed the data: MAF MLD MAI CCN. Contributed reagents/materials/analysis tools: MRE ELH JJS NVR CCN. Wrote the paper: MAF CCN.