About the Authors

Diana S. Smith

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Kejun Guo

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Bradley S. Barrett

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Karl J. Heilman

Affiliation Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America

Leonard H. Evans

Affiliation Rocky Mountain Laboratories, National Institutes of Allergy and Infectious Diseases, Hamilton, Montana, United States of America

Kim J. Hasenkrug

Affiliation Rocky Mountain Laboratories, National Institutes of Allergy and Infectious Diseases, Hamilton, Montana, United States of America

Warner C. Greene

Affiliations Gladstone Institute of Virology and Immunology, University of California San Francisco, San Francisco, California, United States of America, Departments of Medicine, Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Mario L. Santiago

mario.santiago@ucdenver.edu

Affiliations Department of Medicine, University of Colorado Denver, Aurora, Colorado, United States of America, Department of Immunology, University of Colorado Denver, Aurora, Colorado, United States of America, Department of Microbiology, University of Colorado Denver, Aurora, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: M. Santiago, W. Greene, K. Hasenkrug, D. Smith. Performed the experiments: D. Smith, K. Guo, B. Barrett, K. Heilman, L. Evans, K. Hasenkrug, M. Santiago. Analyzed the data: M. Santiago, D. Smith, K. Guo, B. Barrett, L. Evans, K. Hasenkrug. Wrote the paper: M. Santiago, D. Smith, K. Hasenkrug, L. Evans, W. Greene.