About the Authors

Juan Qian

Contributed equally to this work with: Juan Qian, Hui Zheng

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Hui Zheng

Contributed equally to this work with: Juan Qian, Hui Zheng

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Wei-Chun HuangFu

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Jianghuai Liu

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Christopher J. Carbone

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

N. Adrian Leu

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Darren P. Baker

Affiliation BiogenIdec, Cambridge, Massachusetts, United States of America

Serge Y. Fuchs

syfuchs@vet.upenn.edu

Affiliation Department of Animal Biology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

Authors declare no COI except for D.P. Baker, who is employed by BiogenIdec, Inc., and owns the stock.

Author Contributions

Conceived and designed the experiments: JQ WH JL SYF. Performed the experiments: JQ HZ WH JL CJC NAL. Analyzed the data: JQ HZ DPB SYF. Contributed reagents/materials/analysis tools: DPB. Wrote the paper: JQ DPB SYF.