About the Authors

Yorifumi Satou

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Jun-ichirou Yasunaga

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Tiejun Zhao

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Mika Yoshida

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Paola Miyazato

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Ken Takai

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Kei Shimizu

Affiliation Department of Pathology, Kurume University School of Medicine, Kurume, Japan

Koichi Ohshima

Affiliation Department of Pathology, Kurume University School of Medicine, Kurume, Japan

Patrick L. Green

Affiliation Center for Retrovirus Research and Departments of Veterinary Biosciences and Molecular Virology, Immunology and Medical Genetics, The Ohio State University, Columbus, Ohio, United States of America

Naganari Ohkura

Affiliation Department of Experimental Pathology, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Tomoyuki Yamaguchi

Affiliation Department of Experimental Pathology, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Masahiro Ono

Affiliation Department of Experimental Pathology, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Shimon Sakaguchi

Affiliation Department of Experimental Pathology, Institute for Frontier Medical Sciences, Kyoto University, Kyoto, Japan

Masao Matsuoka

mmatsuok@virus.kyoto-u.ac.jp

Affiliation Laboratory of Virus Control, Institute for Virus Research, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YS JY MM. Performed the experiments: YS JY TZ MY PM KT KS KO. Analyzed the data: YS KO NO SS MM. Contributed reagents/materials/analysis tools: PLG TY MO. Wrote the paper: YS JY PLG NO SS MM.