About the Authors

Hilo Yen

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Tadasuke Ooka

Affiliation Division of Microbiology, Department of Infectious Diseases, Faculty of Medicine, Frontier Science Research Center, University of Miyazaki, Miyazaki, Japan

Atsushi Iguchi

Affiliation Interdisciplinary Research Organization, Frontier Science Research Center, University of Miyazaki, Miyazaki, Japan

Tetsuya Hayashi

Affiliations Division of Microbiology, Department of Infectious Diseases, Faculty of Medicine, Frontier Science Research Center, University of Miyazaki, Miyazaki, Japan, Division of Bioenvironmental Science, Frontier Science Research Center, University of Miyazaki, Miyazaki, Japan

Nakaba Sugimoto

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Toru Tobe

torutobe@bact.med.osaka-u.ac.jp

Affiliation Department of Microbiology and Immunology, Graduate School of Medicine, Osaka University, Suita, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HY TH TT. Performed the experiments: HY. Analyzed the data: HY TT. Contributed reagents/materials/analysis tools: TO AI TH NS. Wrote the paper: HY TT.