About the Authors

Gráinne H. Long

grainne.long@lshtm.ac.uk

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Veterinary and Biomedical Sciences, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Divya Sinha

Affiliation Veterinary and Biomedical Sciences, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Andrew F. Read

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Stacy Pritt

Affiliation Covance Research Products Incorporated, Denver, Pennsylvania, United States of America

Barry Kline

Affiliation Covance Research Products Incorporated, Denver, Pennsylvania, United States of America

Eric T. Harvill

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Veterinary and Biomedical Sciences, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Peter J. Hudson

Affiliation Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America

Ottar N. Bjørnstad

Affiliations Center for Infectious Disease Dynamics, Department of Biology, The Pennsylvania State University, University Park, Pennsylvania, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ETH. Performed the experiments: GHL DS SP. Analyzed the data: GHL AFR PJH ONB. Contributed reagents/materials/analysis tools: GHL SP BK ETH PJH ONB. Wrote the paper: GHL AFR PJH ONB.