About the Authors

Ram Samudrala

Affiliation Department of Microbiology, University of Washington, Seattle, Washington, United States of America

Fred Heffron

Affiliation Department of Molecular Microbiology and Immunology, Oregon Health and Science University, Portland, Oregon, United States of America

Jason E. McDermott

Jason.McDermott@pnl.gov

Affiliation Computational Biology and Bioinformatics, Pacific Northwest National Laboratory, Richland, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RS FH JEM. Performed the experiments: JEM. Analyzed the data: JEM. Contributed reagents/materials/analysis tools: RS FH JEM. Wrote the paper: JEM.