About the Authors

Naotoshi Tsuji

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Takeharu Miyoshi

Affiliation Laboratory of Parasitic Diseases, National Institute of Animal Health, National Agriculture and Food Research Organization, Tsukuba, Ibaraki, Japan

Badger Battsetseg

Affiliation National Research Center for Protozoan Diseases, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan

Tomohide Matsuo

Affiliations National Research Center for Protozoan Diseases, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan, Department of Infectious Diseases, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Xuenan Xuan

Affiliation Department of Infectious Diseases, Kyorin University School of Medicine, Mitaka, Tokyo, Japan

Kozo Fujisaki

tick@ms.kagoshima-u.ac.jp

Affiliations National Research Center for Protozoan Diseases, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Hokkaido, Japan, Department of Emerging Infectious Diseases, School of Veterinary Medicine, Kagoshima University, Korimoto, Kagoshima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: N. Tsuji, K. Fujisaki. Performed the experiments: T. Miyoshi, B. Battsetseg, T. Matsuo, X. Xuan. Analyzed the data: N. Tsuji, X. Xuan, K. Fujisaki. Contributed reagents/materials/analysis tools: B. Battsetseg, T. Matsuo. Wrote the paper: N. Tsuji, K. Fujisaki.