About the Authors

Panisadee Avirutnan

Affiliations Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America , Medical Molecular Biology Unit, Office for Research and Development, Mahidol University, Bangkok, Thailand

Lijuan Zhang

Affiliation Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Nuntaya Punyadee

Affiliation Medical Molecular Biology Unit, Office for Research and Development, Mahidol University, Bangkok, Thailand

Ananya Manuyakorn

Affiliation Department of Pathology, Faculty of Medicine Siriraj Hospital, Mahidol University, Bangkok, Thailand

Chunya Puttikhunt

Affiliation Medical Biotechnology Unit, National Center for Genetic Engineering and Biotechnology BIOTEC, National Science and Technology Development Agency NSTDA, Pathumthani, Thailand

Watchara Kasinrerk

Affiliations Medical Biotechnology Unit, National Center for Genetic Engineering and Biotechnology BIOTEC, National Science and Technology Development Agency NSTDA, Pathumthani, Thailand , Department of Medical Technology, Faculty of Associated Medical Sciences, Chiang Mai University, Chiang Mai, Thailand

Prida Malasit

Affiliation Medical Biotechnology Unit, National Center for Genetic Engineering and Biotechnology BIOTEC, National Science and Technology Development Agency NSTDA, Pathumthani, Thailand

John P Atkinson

Affiliations Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America , Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America , Department of Molecular Microbiology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Michael S Diamond

To whom correspondence should be addressed. E-mail: diamond@borcim.wustl.edu

Affiliations Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America , Department of Pathology and Immunology, Washington University School of Medicine, St. Louis, Missouri, United States of America , Department of Molecular Microbiology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

PA and LZ conceived, designed, and performed experiments. NP, AM, CP, WK, PM, JPA, and MSD conceived and designed experiments. PA, LZ, PM, JPA, and MSD wrote the paper.