About the Authors

John Carroll

Contributed equally to this work with: John Carroll, Dilip Dey, Lori Kreisman

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Dilip Dey

Contributed equally to this work with: John Carroll, Dilip Dey, Lori Kreisman

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Lori Kreisman

Contributed equally to this work with: John Carroll, Dilip Dey, Lori Kreisman

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Palanivel Velupillai

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Jean Dahl

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Samuel Telford

Affiliation Department of Tropical Public Health, Harvard School of Public Health, Boston, Massachusetts, United States of America

Roderick Bronson

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Thomas Benjamin

To whom correspondence should be addressed. E-mail: thomas_benjamin@hms.harvard.edu

Affiliation Department of Pathology, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

TB conceived and designed the experiments. JC, DD, LK, and PV performed the experiments. JD, RB, and TB analyzed the data. JC and ST contributed reagents/materials/analysis tools. TB wrote the paper.