About the Authors

Justin Widener

Affiliations Program in Pathobiology, Brown University, Providence, Rhode Island, United States of America , Department of Biochemistry and Molecular Biology, University of Georgia, Athens, Georgia, United States of America

Marianne Jensby Nielsen

Affiliation Department of Medical Biochemistry, University of Aarhus, Aarhus, Denmark

April Shiflett

Affiliation Department of Biochemistry and Molecular Biology, University of Georgia, Athens, Georgia, United States of America

Søren Kragh Moestrup

Affiliation Department of Medical Biochemistry, University of Aarhus, Aarhus, Denmark

Stephen Hajduk

To whom correspondence should be addressed. E-mail: shajduk@bmb.uga.edu

Affiliation Department of Biochemistry and Molecular Biology, University of Georgia, Athens, Georgia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JW, MJN, AS, SKM, and SH conceived and designed the experiments. JW, MJN, and AS performed the experiments and analyzed the data. JW and MJN contributed reagents/materials/analysis tools. JW, AS, SKM, and SH wrote the paper.