About the Authors

Art F. Y Poon

To whom correspondence should be addressed. E-mail: afpoon@ucsd.edu

Affiliation Department of Pathology, University of California San Diego, La Jolla, California, United States of America

Sergei L. Kosakovsky Pond

Affiliation Department of Pathology, University of California San Diego, La Jolla, California, United States of America

Phil Bennett

Affiliation Science Park, University of Warwick, Coventry, United Kingdom

Douglas D Richman

Affiliations Department of Pathology, University of California San Diego, La Jolla, California, United States of America, School of Medicine, University of California San Diego, La Jolla, California, United States of America, Veterans Affairs San Diego Healthcare System, San Diego, California, United States of America

Andrew J. Leigh Brown

Affiliation Institute of Evolutionary Biology, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Simon D. W Frost

Affiliation Department of Pathology, University of California San Diego, La Jolla, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

DDR, AJLB, and SDWF conceived and designed the experiments. AFYP performed the experiments and wrote the paper. AFYP and SLKP analyzed the data. AFYP, SLKP, and PB contributed reagents/materials/analysis tools.