About the Authors

Alex Toftgaard Nielsen

Contributed equally to this work with: Alex Toftgaard Nielsen, Nadia A Dolganov

Affiliation Division of Infectious Diseases and Geographic Medicine, Department of Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Nadia A Dolganov

Contributed equally to this work with: Alex Toftgaard Nielsen, Nadia A Dolganov

Affiliation Division of Infectious Diseases and Geographic Medicine, Department of Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Glen Otto

Affiliations Department of Comparative Medicine, Stanford University, Stanford, California, United States of America , Section of Molecular Genetics and Microbiology, University of Texas at Austin, Austin, Texas, United States of America

Michael C Miller

Affiliation Division of Infectious Diseases and Geographic Medicine, Department of Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Cheng Yen Wu

Affiliation Division of Infectious Diseases and Geographic Medicine, Department of Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Gary K Schoolnik

To whom correspondence should be addressed. E-mail: schoolni@cmgm.stanford.edu

Affiliation Division of Infectious Diseases and Geographic Medicine, Department of Medicine, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

ATN, NAD, and GKS conceived and designed the experiments. ATN, NAD, and GO performed the experiments. ATN and NAD analyzed the data. GO performed all rabbit operations. ATN, NAD, MCM, and CYW contributed reagents/materials/analysis tools. ATN and GKS wrote the paper.